Some types of fill areas have an origin, marked by a orange circle 
. This is used to move the center point for Radial fill, Spiral fill and Shape fill types, or the start point for the pattern in a Pattern Fill. For Pattern Fill, the origin marker also has an adjustment handle to set the angle of the pattern 
.
This means you can control how the pattern is repeated in a pattern fill area.
For Radial fill, Spiral fill and Shape fill areas, the origin can be moved. This sets the start point for the fill, such as the point from which stitches radiate on a Radial fill.
Use Align Origin Handles 
to align the origin for multiple fill areas.